Secreted . Cytoplasmic vesicle, secretory vesicle, acrosome . Cytoplasmic vesicle, secretory vesicle, acrosome lumen . Localized in perinuclear pro-acrosomal granules in round spermatides. Localized between the inner and outer acrosomal membranes (matrix or lumen) in spermatozoa. Secreted during acrosome exocytosis. .
Expressed in spermatozoa (at protein level) (PubMed:22495889).